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Beloved Galápagos tortoise ‘Gramma’ dies at 141

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Beloved Galápagos tortoise ‘Gramma’ dies at 141

December 1, 2025

A member of the first batch of Galápagos tortoises to arrive at the San Diego Zoo over a century ago died last week, the zoo announced Monday. Her age at death was estimated at 141, making her the oldest living animal at the San Diego Zoo.

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